Graduate Seminar Courses

For more information on courses, visit the Evolution and Ecology course distinctions page.
Spring 2019
EEOB 5194
Group Studies: Evolutionary Ecology
Instructors: Ian Hamilton, and Gerry Carter,
Meeting Time: Tuesdays and Thursdays 9:35AM-10:55AM
Course overview
This course uses evolutionary approaches to understand the causes and consequences of interactions between organisms and their environment, including interactions within and between species.  Key topics in the field of evolutionary ecology include the evolution of schedules of reproduction and development (life history), the evolution of behavior and the evolution of interactions among species.
Course Objectives
·Students will understand the diversity of life history patterns, behaviors and interspecific interactions, and how these can be the result of natural, sexual and social selection
·Students will be able to develop and test hypotheses about the evolutionary causes and consequences of variation in life history, behavior and interspecific interactions.
·Students will be able to use mathematical and computational models such as quantitative genetics models and game theoretical models, to make predictions about the evolution of strategies
·Students will understand how evolutionary perspectives are used to understand how individuals, populations and species respond to anthropogenic change.
EEOB 5797
Tropical Behavioral Ecology and Evolution in Panama
Instructors: Rachelle Adams, and Gerry Carter,
Tropical Behavioral Ecology and Evolution in Panama is a May study abroad program where you will be immersed in biological research in a Panamanian Neotropical rainforest. With guidance of instructors, students will propose and complete independent research projects related to evolution, ecology and/or behavior. You will have unique opportunities to interact with a global community of scientists and work at the world-renowned Smithsonian Tropical Research Institute. Funding information can be found at the Office of International Affairs.  For questions contact OSU study abroad ( or Dr. Note: All students must take Undergraduate Independent Studies EEOB 3193 or EEOB Graduate Seminar: Behavior EEOB 8896.02 with Dr. Adams.
EEOB 6330
Phylogenetic Methods
Instructors: John Freudenstein, and Laura Kubatko,
4 credit hours
Class# 33909
Methods for phylogenetic analysis: cladistics, maximum likelihood, Bayesian approaches; character coding and data management.
EEOB 8896.01 
Taste and smell in arthropods
Instructor: Norm Johnson,
14 Weeks; 1 credit hour
Course #34376
A review and discussion of recent literature on the molecular biology, physiology, neurobiology of olfaction and gustation in arthropods. 
EEOB 8896.02
Tropical Research Proposal Writing
Instructors: Rachelle Adams, and Gerry Carter,
We will focus on evolutionary processes that shape the ecology and behavior of systems in a diverse tropical forest. Students will build skills in writing and experimental design in Columbus, Ohio (SP19) as they prepare their research permits and research proposals. Field research will be conducted at the world renowned Smithsonian Tropical Research Institute in Panama (May 2019; EEOB5798). In addition, the students will review and discuss the work of their peers as well as attend lectures to prepare for fieldwork. A final proposal will be submitted in April 2017 in the format of a NSF Preproposal and STRI Short Term Fellowship. This course provides unique opportunities to interact with a global community of scientists and learn successful research strategies while preparing to work in a Neotropical rainforest.
**This course is offered only in conjunction with EEOB 5798
Global biodiversity and macroecology
Instructor: Marta Jarzyna,
Second 7 weeks
Meeting times TBA
Class# 34814
Most contemporary empirical ecology focuses on the ecological interactions of individuals and populations at small temporal and spatial scales and often relies on experimental manipulation. Macroecology is a rapidly emerging field that takes a much larger perspective and explores patterns and processes driving biodiversity across the landscape, continental, and global spatial scales as well as long—i.e., decadal to paleontological—time scales. Because experimental manipulation across such large scales is impossible, macroecology often relies on sophisticated statistical methods. The objective of the course is to introduce students to the ecological questions that are at the forefront of the field of macroecology and to the variety of statistical approaches for tackling them.
EEOB 8896.09
Instructor: Roman Lanno,
Schedule TBA; 2 credit hours 
Class# 33895
EEOB 8896.12
Introduction to Methods in Genome Analysis
Instructors: Alexander Ochoa,; Lisle Gibbs,; Michael Broe,
Second 7 week session
Meeting Time: Wednesdays 10:00AM-12:00PM, 104 Aronoff Laboratory
Class# 33503
EEOB 8896.15
Topics in Population Genetics and Phylogenetics
Instructor: Laura Kubatko, and Paul Fuerst,
Meeting Time: Mondays 12PM-1PM, Aronoff Laboratory
14 Weeks; 1 credit hour
Class# 28427
In this course, we will be reading and discussing the recently published book "Phylogenetic Comparative Methods: Learning from Trees" by Luke Harmon. The phylogenetic comparative method seeks to use a phylogenetic framework to study the evolution of traits ranging from morphological features of organisms to rates of speciation and extinction. The methods used for these analyses include a variety of mathematical and statistical techniques, and our goal will be to achieve an intuitive understanding of how these tools are used to address evolutionary questions of interest. We will read approximately one chapter per week and discuss the reading as a group.
EEOB 8896.19
Quantitative Methods: Multivariate Statistics  
Instructor: Dr. Stuart Ludsin,, Ben Marcek,
2 credit hours
Meeting Time: TBD, 0124 Aquatic Ecology Lab/Research Center
Class# 34984


Micro 5161
Bioinformatics and Genomics
Instructor: Dr. Igor Jouline,, Department of Microbiology
Meeting Time: Monday, Wednesday, Friday; time and location TBD; 3 units
In this course, you will learn how the genome sequencing technology has revolutionized biology and provided a foundation for new developments in science and medicine. You will become familiar with computational tools that are necessary to analyze genomic data and you will find out what biological questions can be answered by genomic approaches. We will use prokaryotes as the main material for genomic studies, but the core principles that you will learn are also applicable to eukaryotes including humans.
More information File here.

Autumn 2018

EEOB 8896.04-190
Graduate Writing Seminar I

Instructor: Ian Hamilton,
First 7 week session (Aug. 28 – Oct. 8); 1.5 credits
Meeting Time: Mondays 1:00-3:00, 104 Aronoff Laboratory
Class # 14587

This graduate-level course is the first of a two-part set that will focus on improving the scientific writing ability of students. This course will focus specifically on writing scientific manuscripts. During the course, students will learn how to approach the writing process, practice writing, and learn to effectively critique their own writing and that of others. For more information, download theFile syllabus.
EEOB 8896.04-290
Graduate Writing Seminar II

Instructor: Ian Hamilton,
Second 7 week session (Oct. 15 – Dec. 3); 1.5 credits
Meeting Time: Mondays 1:00-3:00, 104 Aronoff Laboratory
Class # 18469
This graduate-level course is the second of a two-part set that will focus on improving the scientific writing ability of students. This course will focus specifically on writing scientific proposals. During the course, students will learn how to approach the proposal writing process, practice writing, and learn to effectively critique their own writing and that of others. For more information, download theFile syllabus.

EEOB 8896.10
Comparative phylogeography
Instructor: Bryan Carstens, 
14 weeks
Meeting times TBA
Class # 16091

This seminar will survey classic and current approaches to comparative phylogeography, including gene tree concordance, suture zones & breaks, hypothesis testing, hierarchical approximate Bayesian computation, community trees, automated phylogeography, and likelihood based approaches. The course will be a mix of paper discussion and hands-on experience with software packages, and a term project will be required.

Spring 2018

EEOB 8896.10
The practice of taxonomy and nomenclature

Instructor: Marymegan Daly,

EEOB 8896.11 Graduate Seminar
Effects of climate change on populations and communities
Instructor: Allison Snow,
EEOB 8896.14 Graduate Seminar
Climate change impacts on the structure and function of lake phytoplankton and zooplankton communities.
Instructor: Dr. Jim Hood,
EEOB 8896.15
Seminar in Population Genetics
Instructors: Laura Kubatko,; Paul Fuerst,


Autumn 2017
EEOB 8896.04
Writing Science: Two-part Graduate Seminar Series
Instructor: Stuart Ludsin,
EEOB 8896.07
Communicating science via photography

Instructor: Andi Wolfe,

May / Summer Semester 2017

EEOB 5798
Tropical Behavioral Ecology and Evolution in Panama
Instructor: Rachelle Adams,
EEOB 7210
Methods in Evolution and Ecology: Essential tools for computational biology

Instructors: Bryan Carstens, and Ariadna Morales,

Spring 2017

EEOB 8896.01 

The Anatomy and Physiology of Host-Microbial Interactions
Instructor: Zakee Sabree,

EEOB 8896.02
Graduate Seminar: Behavior
Instructor: Rachelle Adams,
EEOB 8896.05  
Ecological Speciation
Instructor: Bryan Carstens,
EEOB 8896.11
Population Ecology Part II: developing and applying demographic models 
Instructors: Maria Miriti ( and Libby Marschall (
EEOB 8896.18
Sexual Selection and…  
Instructors: Susan Gershman ( and Andy Roberts (
EEOB 8896.19
Agent Based Models in Behavior and Ecology
Instructor: Ian Hamilton,

Autumn 2016

EEOB 8896.10 
Discussion of current primary literature and major topics in phylogenetic systematics.
Instructor: Meg Daly

EEOB 8896.12 
When should engineered gene drives be used to manage wild species?

Instructor: Professor Allison Snow,

EEOB 8896.15  
Seminar in Population Genetics

Instructors:  Laura Kubatko and Paul Fuerst
EEOB 8896.19
Meta-Analysis in Evolution and Ecology
Instructor: Steve Hovick

EEOB 8896.20
Gender in Science and Engineering
Instructor: Joan M. Herbers

Spring 2016

EEOB 6210

Instructor: Roman Lanno, Phone: 292-4943

EEOB8896.04—section 310
Publishing Papers in Plant Ecology
Instructor:  Allison Snow

EEOB 8896.05
Molecular Basis of Adaptive Variation in Natural Populations
Instructor: H. Lisle Gibbs

EEOB 8896.07-100
Communicating science via photography

Instructor: Andi Wolfe

EEOB 8896.09
Hydraulic Fracturing: Environmental and sociological considerations

Instructor: Roman Lanno, Phone: 292-4943

Population Genetics

Instructor: Laura Kubatko

Autumn 2015

Ecology of Inland Waters
Jim Bauer

EEOB8896.04 – 500
Writing Science Graduate Seminar
Stuart Ludsin, Michael Fraker, Conor Keitzer

Evolutionary radiations seminar

Andi Wolfe

EEOB 8896.11
Teaching College-level Ecology: Bringing Innovation Into the Laboratory Experience

Maria Miriti and Peter Curtis

Molecular Ecology

Bryan Carstens

Host finding mechanisms in insects

Norm Johnson

Current Topics in Quantative Methods

Bryan Carstens

Spring 2015

EEOB8896.02 – 500
Graduate Seminar in Complex Adaptive Systems in Ecology, Evolution, and Behavior

Dr. Ian Hamilton

EEOB 8896.05
Evolutionary Innovation

Sponsored by John Freudenstein

EEOB 8896.12
Model-based Phylogeography

Dr. Bryan Carstens

EEOB 8896.15
Topics in Phylogenomic Inference

Instructor: Laura Kubatko 

EEOB 8896.20
WGSS 8896.20
Gender in Science and Engineering

Dr. Joan M. Herbers

Autumn 2014

EEOB 5450
Quantitative Population Ecology

Drs. Maria Miriti, Libby Marschall, and Noelle Beckman

EEOB 8896.02
Animal Personalities
Instructor: Dr. Doug Nelson

EEOB 8896.04 
Career Options for EEOB PhDs: Exploring the Landscape
Instructor: Dr. Allison Snow, EEOB

EEOB 8896.04
EEOB Departmental Seminar Series
Zakee Sabree

EEOB 8896.07-100 (34551)
Communicating science via photography
Instructor: Andi Wolfe

EEOB 8896.12
Molecular Ecology focusing on Phylogeography and Landscape genetics

Instructor: Bryan Carstens

Spring 2014

EEOB 5320
Creation and Evolution: Differing Worldviews  (29190)

Instructor: Dr. Andrea D. Wolfe

EEOB 6210

Instructor: Roman Lanno

EEOB 6320
Principles of Systematics I

Instructor: Hans Klompen

EEOB 7220
Modeling in Evolutionary Ecology

Instructors: Ian Hamilton, Libby Marschall, David Glover

EEOB 8896.04
Graduate Seminar in Ecology: Ecology and evolution in weedy and invasive species: considering plant functional traits

Instructors: Steve Hovick and Kristin Mercer (HCS faculty)

EEOB 8896.05
Evolution and ecology of animal venoms: Unanswered questions

Instructors: Meg Daly, Lisle Gibbs

EEOB 8896.06
Phylogeography and evolution in marine systems

Instructor: Meg Daly

EEOB 8896.15
Current Topics in Mathematical Population Genetics

Instructors:  Laura Kubatko and Paul Fuerst



The Nature and Practice of Science

Dr. Dan Herms ,  Professor and Chair, Dept. of Entomology
Dr. Carol Anelli, Professor and Associate Chair, Dept. of Entomology                                          
Dr. Ross MacDonald, Research Scientist, Dept. of Entomology

Commercialization for Researchers

Team Leader: Professor Thomas Rosol, 614-292-5661

Autumn 2013

EEOB 5420
Ecology of Inland Waters
Instructor: Dr. James Bauer

EEOB 7310
Studies in Taxonomy (to become Plant Taxonomy)

Instructor: Dr. John Freudenstein
This offering of the course will focus on vascular plant taxonomy.

EEOB 8896.02 
Behavior Seminar

Instructors: Dr. Andrew Roberts & Dr. Susan Gershman
Animal Behavior Seminar

EEOB 8896.04
Publishing in High-Impact Journals
Instructor: Dr. Allison Snow

EEOB 8896.10
Graduate Seminar in Systematics:

Instructor: Bryan Carstens

EEOB 8896.11
Foundations of Quantitative Ecology Seminar

Instructor: Paul Hurtado

EEOB 8896.11
Graduate Seminar in Population Ecology:
Introduction to R for Biologists

Instructor: Simon Queenborough