Graduate Seminar Courses

Spring 2019
EEOB 5194
Group Studies: Evolutionary Ecology
Instructors: Ian Hamilton, and Gerry Carter,
Meeting Time: Tuesdays and Thursdays 9:35AM-10:55AM
Course overview
This course uses evolutionary approaches to understand the causes and consequences of interactions between organisms and their environment, including interactions within and between species.  Key topics in the field of evolutionary ecology include the evolution of schedules of reproduction and development (life history), the evolution of behavior and the evolution of interactions among species.
Course Objectives
·Students will understand the diversity of life history patterns, behaviors and interspecific interactions, and how these can be the result of natural, sexual and social selection
·Students will be able to develop and test hypotheses about the evolutionary causes and consequences of variation in life history, behavior and interspecific interactions.
·Students will be able to use mathematical and computational models such as quantitative genetics models and game theoretical models, to make predictions about the evolution of strategies
·Students will understand how evolutionary perspectives are used to understand how individuals, populations and species respond to anthropogenic change.
EEOB 5797
Tropical Behavioral Ecology and Evolution in Panama
Instructors: Rachelle Adams, and Gerry Carter,
Tropical Behavioral Ecology and Evolution in Panama is a May study abroad program where you will be immersed in biological research in a Panamanian Neotropical rainforest. With guidance of instructors, students will propose and complete independent research projects related to evolution, ecology and/or behavior. You will have unique opportunities to interact with a global community of scientists and work at the world-renowned Smithsonian Tropical Research Institute. Funding information can be found at the Office of International Affairs.  For questions contact OSU study abroad ( or Dr. Note: All students must take Undergraduate Independent Studies EEOB 3193 or EEOB Graduate Seminar: Behavior EEOB 8896.02 with Dr. Adams.
EEOB 6330
Phylogenetic Methods
Instructors: John Freudenstein, and Laura Kubatko,
4 credit hours
Class# 33909
Methods for phylogenetic analysis: cladistics, maximum likelihood, Bayesian approaches; character coding and data management.
EEOB 8896.02
Graduate Seminar: (Behavior)—Tropical Research Proposal Writing
Instructors: Rachelle Adams, and Gerry Carter,
We will focus on evolutionary processes that shape the ecology and behavior of systems in a diverse tropical forest. Students will build skills in writing and experimental design in Columbus, Ohio (SP19) as they prepare their research permits and research proposals. Field research will be conducted at the world renowned Smithsonian Tropical Research Institute in Panama (May 2019; EEOB5798). In addition, the students will review and discuss the work of their peers as well as attend lectures to prepare for fieldwork. A final proposal will be submitted in April 2017 in the format of a NSF Preproposal and STRI Short Term Fellowship. This course provides unique opportunities to interact with a global community of scientists and learn successful research strategies while preparing to work in a Neotropical rainforest.
**This course is offered only in conjunction with EEOB 5798
EEOB 8896.09
Instructor: Roman Lanno,
Schedule TBA; 2 credit hours 
Class# 33895
EEOB 8896.12
Introduction to Methods in Genome Analysis
Instructors: Alexander Ochoa,; Lisle Gibbs,; Michael Broe,
Second 7 week session
Meeting Time: Wednesdays 10:00AM-12:00PM, 104 Aronoff Laboratory
Class# 33503
EEOB 8896.15
Instructor: Laura Kubatko, and Paul Fuerst,


Micro 5161
Bioinformatics and Genomics
Instructor: Dr. Igor Jouline,, Department of Microbiology
Meeting Time: Monday, Wednesday, Friday; time and location TBD; 3 units
In this course, you will learn how the genome sequencing technology has revolutionized biology and provided a foundation for new developments in science and medicine. You will become familiar with computational tools that are necessary to analyze genomic data and you will find out what biological questions can be answered by genomic approaches. We will use prokaryotes as the main material for genomic studies, but the core principles that you will learn are also applicable to eukaryotes including humans.
More information File here.

Autumn 2018

EEOB 8896.04-190
Graduate Writing Seminar I

Instructor: Ian Hamilton,
First 7 week session (Aug. 28 – Oct. 8); 1.5 credits
Meeting Time: Mondays 1:00-3:00, 104 Aronoff Laboratory
Class # 14587

This graduate-level course is the first of a two-part set that will focus on improving the scientific writing ability of students. This course will focus specifically on writing scientific manuscripts. During the course, students will learn how to approach the writing process, practice writing, and learn to effectively critique their own writing and that of others. For more information, download theFile syllabus.
EEOB 8896.04-290
Graduate Writing Seminar II

Instructor: Ian Hamilton,
Second 7 week session (Oct. 15 – Dec. 3); 1.5 credits
Meeting Time: Mondays 1:00-3:00, 104 Aronoff Laboratory
Class # 18469
This graduate-level course is the second of a two-part set that will focus on improving the scientific writing ability of students. This course will focus specifically on writing scientific proposals. During the course, students will learn how to approach the proposal writing process, practice writing, and learn to effectively critique their own writing and that of others. For more information, download theFile syllabus.

EEOB 8896.10
Comparative phylogeography
Instructor: Bryan Carstens, 
14 weeks
Meeting times TBA
Class # 16091

This seminar will survey classic and current approaches to comparative phylogeography, including gene tree concordance, suture zones & breaks, hypothesis testing, hierarchical approximate Bayesian computation, community trees, automated phylogeography, and likelihood based approaches. The course will be a mix of paper discussion and hands-on experience with software packages, and a term project will be required.

Spring 2018

EEOB 8896.10
The practice of taxonomy and nomenclature

Instructor: Marymegan Daly,
14 weeks; 1 credit 
Meeting times TBA; One 2-hour meeting/week
Class # 36996

Course Learning Objectives
Recognize and understand the conceptual and methodological issues that inform the delimitation of taxa.
Recognize and apply rules of nomenclature that pertain to the description and synonymy of taxa.
Use written and oral communication to critique and interpret taxonomic and nomenclatural acts.
Understand the intersection of nomenclature, systematics, and conservation 
EEOB 8896.11 Graduate Seminar
Effects of climate change on populations and communities
Instructor: Allison Snow,
Second 7-week session (March 2 – April 20), 1 credit
Meeting Time: Friday 11:00AM-1:00PM, Aronoff Lab 104
Course #36037
In this graduate-level seminar, we will examine the extent to which ongoing climate change has affected the distribution, phenology, life history stages, and ecological interactions of various species.  Each week, we will discuss peer-reviewed publications to examine methods, evidence, and conclusions about these ecological signatures of climate change.  Our primary goals are to become familiar with some of the strongest evidence for how wild species already have been affected by directional changes in climatic conditions, and to evaluate related research on possible future effects.  For example, which phenological and life history shifts in plant or animal populations have been linked to global warming?  How has climate change affected interactions among species, such as mutualisms, competition, and food webs?  What is the evidence for climate change leading to population declines, niche shifts, or range shifts, and why are some species affected more than others?  Readings will be selected from top journals such as PNAS, Science, Nature, Ecology Letters, and others, and topics will be chosen to reflect participants’ study systems when possible. 
EEOB 8896.14 Graduate Seminar
Climate change impacts on the structure and function of lake phytoplankton and zooplankton communities.
Instructor: Dr. Jim Hood,
14 weeks; 1 credit
Meeting time and place: TBD based on participants’ schedules. If you plan to enroll in this seminar, contact Jim Hood ( soon so we can schedule a meeting time.
Course #36322 
In this graduate seminar, we will examine how climate change influences the structure and function freshwater phytoplankton and zooplankton communities. Our primary goal will be to understand how climate change influences the physiochemical environment in lakes (i.e., ice cover, water temperatures, physical mixing, nutrient and solute loading, etc.) and how those changes influence plankton community composition, interactions, and phenology. Throughout this graduate seminar, we will address multiple emerging questions related to climate change impacts on freshwater plankton. For instance, we will ask:  How the timing of warming (e.g., winter vs. summer) influences plankton succession and community structure? Whether climate change enhances the severity of harmful algal blooms? And how climate change influences the stability of plankton communities? To address these questions, we will discuss recent peer-reviewed publications concerning the impacts of climate change on plankton communities paired, when appropriate, with related foundational publications. The specific questions and publications will be tailored to match participates’ interests. 
Plankton & Climate Change Seminar
EEOB 8896.15
Seminar in Population Genetics
Instructors: Laura Kubatko,; Paul Fuerst,
1 credit hour
Class # 34221
Meeting: TBD
In this seminar, we will study the statistical methodology used to model population genetics and phylogenetic processes in a coalescent framework. For Spring Semester 2018, we will focus on modeling gene flow and other processes that generate reticulate evolution, such as recombination.


Autumn 2017
EEOB 8896.04
Writing Science: Two-part Graduate Seminar Series
Instructor: Stuart Ludsin,
First 7 week session (Aug. 22 – Oct. 9, 2017); 1 credit
one 2-hour meeting**
class #22749
Second 7 week session (Oct. 16 – Dec. 6, 2017); 1 credit
one 2-hour meeting**
class #36106
The central goal of this seminar series is to help students improve their ability to write science-based manuscripts and research proposals. The seminars will be independent of one another in the sense that students can take one or both seminars and benefit, with minimal overlap of course material. The general structure of both seminars is outlined below. I will use a University Center for the Advancement of Teaching (UCAT) course-design workshop that I am taking in June 2017 to help finalize the syllabus and approach for both seminars so as to maximize student learning.
Mechanics of Manuscript Writing: 1st 7 weeks of Autumn Semester
The first 7-week course will focus on the nuts and bolts of effective scientific writing (e.g., style, structure/organization, paragraph/sentence development, syntax/word choice), with the aim of teaching how to develop coherent, interesting papers that will get cited. While the focus will be on manuscript writing, the lessons learned will also be applicable to proposal writing. We will use one or both of the following textbooks in the course: Joshua Schimel’s Writing Science: How to Write Papers That Get Cited and Proposals That Get Funded (ISBN: 9780199760244) and/or Stephen Heard’s The Scientist’s Guide to Writing: How to Write More Easily and Effectively throughout Your Scientific Career (ISBN: 9781400881147). These two texts, combined with a handful of scientific papers to use as examples, will serve as the foundation for the weekly, student-led discussions. No “major” writing (e.g., manuscript, proposal) would be required for this course. Students would instead be required to help lead a discussion period, complete weekly readings and homework assignments (e.g., editing and critiquing papers; short writing assignments), and attend/participate in class discussions.
Mechanics of Proposal Writing: 2nd 7 weeks of Autumn Semester
Image result for nsf guide to writing proposalsThe second 7-week course will focus more explicitly on developing an effective research proposal. This course will help provide a cookbook approach to writing all components of a research proposal, using Stephen Russell’s and David Morrison’s The Grant Application Writer’s Workbook as a guide. Additional readings (e.g., proposals, blogs, and peer-reviewed literature) will be used as needed. The course will be geared toward developing effective NSF-style proposals; however, the lessons learned will benefit efforts to write any kind of proposal (both long and short), as well as manuscripts. Students will be required to identify a writing goal for the semester, and be expected to write each week toward that goal. Ideally, the writing goal would be to develop (or improve upon) a research proposal (e.g., NSF DDIG, GRF, or pre-proposal; candidacy exam or thesis proposal; small grant application). I would, however, consider allowing a manuscript to be the writing goal, if the student took my previous (1st 7-week) seminar. Students will also be expected to peer-review someone else’s work each week. Each weekly class meeting will likely consist of a short discussion about that week’s readings, followed by an exchange of information about the peer reviews. A commitment to writing and peer-reviewing will be needed to pass this course.  
**Enrolled students will be expected to complete a poll during early summer, to identify a meeting place, day, and time for both courses. Students in the first 7-week course also will be solicited to identify and then read a handful of peer-reviewed papers in advance of the first class meeting, which will facilitate class discussions. The location, time, and day of the seminar will be determined on a later date (May or June), in consultation with enrolled students. Doing so will allow instructor to accommodate as many students as possible, with the recognition that some students inevitably might not be able to take the course.
EEOB 8896.07
Communicating science via photography

Instructor: Andi Wolfe,
First 7-week session; 2 credits
Class #23488

Every scientist needs to communicate his/her research in as many ways as possible. Sharing images during presentations at meetings, while giving seminars, teaching courses, or via social media sites are good ways to let others know about your work and passion(s) for evolution, ecology, or organismal biology.

This seven-week workshop will focus (pun intended) on all aspects of nature photography – from the basics of how to make photos under a variety of conditions to using software for post-processing of images. We’ll meet twice a week – for a 2-hr period during the regular week, and again for a half-day or all-day field trip on Saturday or Sunday to go make photos in local parks, nature reserves, or other place with opportunities for nature photography. Students will need a copy of Lightroom CC (available at Wired Out for an academic discount), and own, rent, or borrow an appropriate camera to use during the course.

Specific goals of the course include: 1) review of photography basics, 2) learn how to use post-processing software, 3) develop a critical eye for composition, exposure, and depth-of-field settings, and 4) sharing photos and information via social media


May / Summer Semester 2017

EEOB 5798
Tropical Behavioral Ecology and Evolution in Panama
Instructor: Rachelle Adams,
EEOB 7210
Methods in Evolution and Ecology: Essential tools for computational biology

Instructors: Bryan Carstens, and Ariadna Morales,

Spring 2017

EEOB 8896.01 

The Anatomy and Physiology of Host-Microbial Interactions
Instructor: Zakee Sabree,

EEOB 8896.02
Graduate Seminar: Behavior
Instructor: Rachelle Adams,
EEOB 8896.05  
Ecological Speciation
Instructor: Bryan Carstens,
EEOB 8896.11
Population Ecology Part II: developing and applying demographic models 
Instructors: Maria Miriti ( and Libby Marschall (
EEOB 8896.18
Sexual Selection and…  
Instructors: Susan Gershman ( and Andy Roberts (
EEOB 8896.19
Agent Based Models in Behavior and Ecology
Instructor: Ian Hamilton,

Autumn 2016

EEOB 8896.10 
Discussion of current primary literature and major topics in phylogenetic systematics.
Instructor: Meg Daly

EEOB 8896.12 
When should engineered gene drives be used to manage wild species?

Instructor: Professor Allison Snow,

EEOB 8896.15  
Seminar in Population Genetics

Instructors:  Laura Kubatko and Paul Fuerst
EEOB 8896.19
Meta-Analysis in Evolution and Ecology
Instructor: Steve Hovick

EEOB 8896.20
Gender in Science and Engineering
Instructor: Joan M. Herbers

Spring 2016

EEOB 6210

Instructor: Roman Lanno, Phone: 292-4943

EEOB8896.04—section 310
Publishing Papers in Plant Ecology
Instructor:  Allison Snow

EEOB 8896.05
Molecular Basis of Adaptive Variation in Natural Populations
Instructor: H. Lisle Gibbs

EEOB 8896.07-100
Communicating science via photography

Instructor: Andi Wolfe

EEOB 8896.09
Hydraulic Fracturing: Environmental and sociological considerations

Instructor: Roman Lanno, Phone: 292-4943

Population Genetics

Instructor: Laura Kubatko

Autumn 2015

Ecology of Inland Waters
Jim Bauer

EEOB8896.04 – 500
Writing Science Graduate Seminar
Stuart Ludsin, Michael Fraker, Conor Keitzer

Evolutionary radiations seminar

Andi Wolfe

EEOB 8896.11
Teaching College-level Ecology: Bringing Innovation Into the Laboratory Experience

Maria Miriti and Peter Curtis

Molecular Ecology

Bryan Carstens

Host finding mechanisms in insects

Norm Johnson

Current Topics in Quantative Methods

Bryan Carstens

Spring 2015

EEOB8896.02 – 500
Graduate Seminar in Complex Adaptive Systems in Ecology, Evolution, and Behavior

Dr. Ian Hamilton

EEOB 8896.05
Evolutionary Innovation

Sponsored by John Freudenstein

EEOB 8896.12
Model-based Phylogeography

Dr. Bryan Carstens

EEOB 8896.15
Topics in Phylogenomic Inference

Instructor: Laura Kubatko 

EEOB 8896.20
WGSS 8896.20
Gender in Science and Engineering

Dr. Joan M. Herbers

Autumn 2014

EEOB 5450
Quantitative Population Ecology

Drs. Maria Miriti, Libby Marschall, and Noelle Beckman

EEOB 8896.02
Animal Personalities
Instructor: Dr. Doug Nelson

EEOB 8896.04 
Career Options for EEOB PhDs: Exploring the Landscape
Instructor: Dr. Allison Snow, EEOB

EEOB 8896.04
EEOB Departmental Seminar Series
Zakee Sabree

EEOB 8896.07-100 (34551)
Communicating science via photography
Instructor: Andi Wolfe

EEOB 8896.12
Molecular Ecology focusing on Phylogeography and Landscape genetics

Instructor: Bryan Carstens

Spring 2014

EEOB 5320
Creation and Evolution: Differing Worldviews  (29190)

Instructor: Dr. Andrea D. Wolfe

EEOB 6210

Instructor: Roman Lanno

EEOB 6320
Principles of Systematics I

Instructor: Hans Klompen

EEOB 7220
Modeling in Evolutionary Ecology

Instructors: Ian Hamilton, Libby Marschall, David Glover

EEOB 8896.04
Graduate Seminar in Ecology: Ecology and evolution in weedy and invasive species: considering plant functional traits

Instructors: Steve Hovick and Kristin Mercer (HCS faculty)

EEOB 8896.05
Evolution and ecology of animal venoms: Unanswered questions

Instructors: Meg Daly, Lisle Gibbs

EEOB 8896.06
Phylogeography and evolution in marine systems

Instructor: Meg Daly

EEOB 8896.15
Current Topics in Mathematical Population Genetics

Instructors:  Laura Kubatko and Paul Fuerst



The Nature and Practice of Science

Dr. Dan Herms ,  Professor and Chair, Dept. of Entomology
Dr. Carol Anelli, Professor and Associate Chair, Dept. of Entomology                                          
Dr. Ross MacDonald, Research Scientist, Dept. of Entomology

Commercialization for Researchers

Team Leader: Professor Thomas Rosol, 614-292-5661

Autumn 2013

EEOB 5420
Ecology of Inland Waters
Instructor: Dr. James Bauer

EEOB 7310
Studies in Taxonomy (to become Plant Taxonomy)

Instructor: Dr. John Freudenstein
This offering of the course will focus on vascular plant taxonomy.

EEOB 8896.02 
Behavior Seminar

Instructors: Dr. Andrew Roberts & Dr. Susan Gershman
Animal Behavior Seminar

EEOB 8896.04
Publishing in High-Impact Journals
Instructor: Dr. Allison Snow

EEOB 8896.10
Graduate Seminar in Systematics:

Instructor: Bryan Carstens

EEOB 8896.11
Foundations of Quantitative Ecology Seminar

Instructor: Paul Hurtado

EEOB 8896.11
Graduate Seminar in Population Ecology:
Introduction to R for Biologists

Instructor: Simon Queenborough