Flowering plant breeding systems, bryophyte ecology. Current foci are: (1) assessing the effects of population structure and pollinator visitations rates on self and outcross pollination in prairie milkweeds, (2) assessing the importance and prevalence of bryophytes in the early establishment of restored wet prairies.  Overall research goals are to better understand the forces influencing plant distribution and plant evolution, and to accomplish environmental enhancement through more effective grassland restoration.